Education
PhD, University of California, Santa Barbara, 2017
MS, University of Maine, 2012
BA, Colby College, 2010
MS, University of Maine, 2012
BA, Colby College, 2010

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Google Scholar
ResearchGate
Please email immccull@gmail.com to obtain manuscript copies if you do not otherwise have access.
Peer-reviewed publications
McCullough, I. M., Brentrup, J. A., Wagner, T., Lapierre J., Henneck, J, Paul, A. M., Belair, M., Moritz, M. A. and C. T. Filstrup. (2023). Fire characteristics and hydrologic connectivity influence short-term responses of north temperate lakes to wildfire. Geophysical Research Letters. https://doi.org/10.1029/2023GL103953
McCullough, I. M., Hanly, P. J., King, K. B. S. and T. Wagner. (2022). Freshwater corridors in the conterminous United States: a coarse-filter approach based on lake-stream networks. Ecosphere. https://doi.org/10.1002/ecs2.4326
DePalma-Dow, A., McCullough, I. M. and J. A. Brentrup. (2022). Turning up the heat: long-term water quality responses to wildfires and climate change in a hypereutrophic lake. Ecosphere. https://doi.org/10.1002/ecs2.4271
Lofton, M., Brentrup, J., Beck, W., Zwart, J., Bhattacharya, R., Brighenti, L., Burnet, S., McCullough, I.M., Steele, B., Carey, C., Cottingham, K., Dietze, M., Ewing, H., Weathers, K. and S. LaDeau. (2022). Using near-term forecasts and uncertainty partitioning to improve predictions of oligotrophic cyanobacterial density. Ecological Applications. https://doi.org/10.1002/eap.2590
Orr, R., Bui, A., Klope, M., McCullough, I. M., Lee, M., Motta, C. Mayorga, I., Konicek, K. and H. Young. Context-dependent effects of shifting large herbivore assemblages on plant structure and diversity. (2022). Journal of Ecology. https://doi.org/10.1111/1365-2745.13871
McCullough, I. M., Fergus., C. E. and C. Scott. Macrosystems limnology and beyond: reenvisioning the scale of limnology. (2022). Encyclopedia of Inland Waters, 2nd edition. ISBN 9780124095489. https://doi.org/10.1016/B978-0-12-819166-8.00026-8
Cheruvelil, K. S., Soranno, P. A., McCullough, I. M., Webster, K. E., Rodriguez, L. K. and N. J. Smith. LAGOS-US-LOCUS v1.0: Data module of location, identifiers, and their watersheds in the conterminous U.S. (2021). Limnology and Oceanography Letters. https://doi.org/10.1002/lol2.10203.
Soranno, P. A., Cheruveli, K. S., Liu, B., Wang, Q, Tan, P., Zhou, J., King, K. B. S., McCullough, I. M., Stachelek, J., Bartley, M., Filstrup, C. T., Hanks, E. M., Lapierre, J., Lottig, N. R., Schliep, E. M., Wagner T. and K. E. Webster. Ecological prediction at macroscales using big data: Does sampling design matter? (2020). Ecological Applications. https://doi.org/10.1002/eap.2123
Wagner, T., Lottig, N., Bartley, M., Hanks, E., Schliep, E., Wikle, N., King, K., McCullough, I. M., Stachelek, J., Cheruvelil, K., Filstrup, C., Lapierre, J., Liu, B., Smith, N., Soranno, P., Tan, P., Wang, Q., Webster, K. and J. Zhou. (2020). Increasing accuracy of lake nutrient predictions in thousands of lakes by leveraging water clarity data. Limnology and Oceanography Letters. https://doi.org/10.1002/lol2.10134
Poisson, A. C., McCullough, I. M., Cheruvelil, K. S., Elliot, K. C., Latimore, J. A. and P. A. Soranno. (2020). Quantifying the contribution of citizen science to broad-scale ecological databases. Frontiers in Ecology and the Environment. https://doi.org/10.1002/fee.2128
McCullough, I. M., Skaff, N. K., Soranno, P. A. and K. S. Cheruvelil. (2019). No lake left behind: how well do US protected areas meet lake conservation targets? Limnology and Oceanography Letters, 4(6): 183-192. https://doi.org/10.1002/lol2.10123
Filstrup, C. T., King, K. B. S. and I. M. McCullough. (2019). Evenness effects mask richness effects on ecosystem functioning at macro-scales in lakes. Ecology Letters, 22(12): 2120-2129. https://doi.org/10.1111/ele.13407
McCullough, I. M., King, K. B. S., Stachelek, J., Diaz, J., Soranno, P. A. and K. S. Cheruvelil. (2019). Applying the patch-matrix model to lakes: a connectivity-based conservation framework. Landscape Ecology, 34(11): 2703-2718. https://doi.org/10.1007/s10980-019-00915-7.
McCullough, I. M., Cheruvelil, K. S., Lapierre, J., Lottig, N. R., Moritz, M. A., Stachelek, J. and P. A. Soranno. (2019). Do lakes feel the burn? Ecological consequences of increasing exposure of lakes to fire in the continental United States. Global Change Biology, 25(9): 2841-2854. https://doi.org/10.1111/gcb.14732.
Davis, F. W., Synes, N. W., Fricker, G. A., McCullough, I. M., Serra-Diaz, S. M., Franklin, J. and A. Flint. (2019). LiDAR-derived topography and forest structure predict fine-scale variation in daily surface temperatures in oak savanna and conifer forest landscapes. Agricultural and Forest Meteorology, 269: 192-202. https://doi.org/10.1016/j.agrformet.2019.02.015.
McCullough, I. M., Cheruvelil, K. S., Collins, S. C. and P. A. Soranno. Geographic patterns of the climate sensitivity of lakes. (2019). Ecological Applications, e01836. https://doi.org/10.1002/eap.1836.
McCullough, I. M., Dugan, H. A., Farrell, K. J., Morales-Williams, A. M., Ouyang, Z., Roberts, D. C., Scordo, F., Bartlett, S. L., Burke, S. M., Doubek, J. P., Krivak-Tetley, F. E., Skaff, N. K., Summers, J. C., Weathers, K. C. and P. C. Hanson. (2018). Dynamic modeling of organic carbon fates in lake ecosystems. Ecological Modelling, 386: 71-82. https://doi.org/10.1016/j.ecolmodel.2018.08.009.
Engel, F., Farrell, K. J., McCullough, I. M., Scordo, F., Denfeld, B. A., Dugan, H. A., de Eyto, E., Hanson, P. C., McClure, R. P., Noges, P., Noges, T., Ryder, E., Weathers, K. C. and G. A. Weyhenmeyer. (2018). A lake classification concept for a more accurate global estimate of the dissolved inorganic carbon export from terrestrial ecosystems to inland waters. The Science of Nature, 105:25. https://doi.org/10.1007/s00114-018-1547-z.
McCullough, I. M., Davis, F. W. and A. P. Williams. (2017). A range of possibilities: assessing geographic variation in climate sensitivity of ponderosa pine using tree-rings. Forest Ecology and Management, 402: 223-233. https://doi.org/10.1016/j.foreco.2017.07.025.
Dugan, H. A., Bartlett, S. L., Burke, S. M., Doubek, J. P., Krivak-Tetley, F. E., Skaff, N. K., Summers, J. C., Farrell, K. J., McCullough, I. M., Morales-Williams, A. M., Roberts, D., Scordo, F., Yang, Z., Hanson, P. C. and K. C. Weathers. (2017). Salting our freshwater lakes. Proceedings of the National Academy of Sciences, 114(17): 4453-4458. https://doi.org/10.1073/pnas.1620211114.
Davis, F. W., Sweet, L., Serra-Diaz, J. M., Franklin, J., McCullough, I. M., Flint, A., Flint, L., Dingman, J. R., Regan, H. M., Syphard, A., Hannah, L., Redmond, K. and M. A. Moritz. (2016). Shrinking windows of opportunity for oak seedling establishment in southern California mountains. Ecosphere, 7(11), article e01573. https://doi.org/10.1002/ecs2.1573.
McCullough, I. M., Davis, F. W., Dingman, J. R., Flint, L. E., Flint, A. L., Serra-Diaz, J. M., Syphard, A. D., Moritz, M. A., Hannah, L. and J. Franklin. (2016). High and dry: high elevations disproportionately exposed to regional climate change in Mediterranean-climate landscapes. Landscape Ecology 31(5): 1063-1075. https://doi.org/10.1007/s10980-015-0318-x.
Serra-Diaz, J. M., Franklin, J., Sweet, L, McCullough, I. M., Syphard, A. D., Regan, H., Flint, L., Flint, A., Dingman, J., Moritz, M. A., Redmond, K., Hannah, L. and F. W. Davis. (2016). Averaged 30 year climate change projections mask opportunities for species establishment. Ecography. 844-845. https://doi.org/10.1111/ecog.02074.
Hannah, L., Flint, L. E., Syphard, A. D., Moritz, M. A., Buckley, L. B. and I. M. McCullough. (2015). Place and process in conservation planning for climate change: a reply to Keppel and Wardell-Johnson. Trends in Ecology and Evolution 30(5): 233-234. https://doi.org/10.1016/j.tree.2015.03.008.
Aceves-Bueno, E., Adeleye, A. S., Bradley, D., Brandt, W. T., Callery, P., Feraud, M., Garner, K. L., Gentry, R., Huang, Y., McCullough, I. M., Pearlman, I., Sutherland, S. A., Wilkinson,W., Yang, Y, Zink, T., Anderson, S. E. and C. Tague. (2015). Citizen science as a tool for overcoming insufficient monitoring and inadequate stakeholder buy-in in adaptive management: Criteria and evidence. Ecosystems 18(3): 493-506. https://doi.org/10.1007/s10021-015-9842-4.
Hannah, L., Flint, L., Syphard, A. D., Moritz, M. A., Buckley, L. B. and I. M. McCullough. (2014). Fine-grain modeling of the response of species to climate change: holdouts, stepping-stones and microrefugia. Trends in Ecology and Evolution 29 (7): 390-397. https://doi.org/10.1016/j.tree.2014.04.006.
Dingman, J. R., Sweet, L. C., McCullough, I. M., Davis, F. W., Flint, A., Franklin, J. and L. E. Flint (2013). Cross-scale modeling of surface temperature and seedling establishment to improve projections of tree distribution shifts under climate change. Ecological Processes 2-30. https://doi.org/10.1186/2192-1709-2-30.
McCullough, I. M., Loftin, C. S. and S. A. Sader. (2013). Landsat imagery reveals declining clarity of Maine’s lakes during 1995-2010. Freshwater Science 32(3): 741-752. https://doi.org/10.1899/12-070.1.
McCullough, I. M., Loftin, C. S. and S. A. Sader. (2013). Lakes without Landsat? An alternative approach to remote lake monitoring with MODIS 250 m imagery. Lake and Reservoir Management 29: 89-98. https://doi.org/10.1080/10402381.2013.778926.
McCullough, I. M., Loftin, C. S. and S. A. Sader. (2012). High-frequency remote monitoring of large lakes with MODIS 500 m imagery. Remote Sensing of Environment 124: 234-241. https://doi.org/10.1016/j.rse.2012.05.018.
McCullough, I. M., Loftin, C. S. and S. A. Sader. (2012). Combining lake and watershed characteristics with Landsat TM data for remote estimation of regional lake clarity. Remote Sensing of Environment 123: 109-115. https://doi.org/10.1016/j.rse.2012.03.006.
Technical reports and other publications
Webster, K. E., McCullough, I. M. and P. A. Soranno. (2022). Deeper by the dozen: Diving into a database of 17,675 depths for U.S. lakes and reservoirs. Limnology and Oceanography Bulletin. https://doi.org/10.1002/lob.10482.
McCullough, I. M., Cheruvelil, K. S., Collins, S. C. and P. A. Soranno. Geographic patterns of the climate sensitivity of lakes. Bulletin of the Ecological Society of America: Photo Gallery. Available here.
Moritz, M.A., Topik, C, Allen, C. D., Hessburg, P. F., Morgan, P., Odion, D. C., Veblen, T. T. and McCullough, I. M. (2018). A Statement of Common Ground Regarding the Role of Wildfire in Forested Landscapes of the Western United States. Fire Research Consensus Working Group Final Report. https://www.nceas.ucsb.edu/featured/fire-research-consensus.
McCullough, I. M., Loftin, C. S. and S. A. Sader. A manual for remote sensing of Maine lake clarity. (2013). Maine Agricultural and Forest Experimentation Station, University of Maine. Technical Bulletin 207. ISSN: 1070-1054.
ResearchGate
Please email immccull@gmail.com to obtain manuscript copies if you do not otherwise have access.
Peer-reviewed publications
McCullough, I. M., Brentrup, J. A., Wagner, T., Lapierre J., Henneck, J, Paul, A. M., Belair, M., Moritz, M. A. and C. T. Filstrup. (2023). Fire characteristics and hydrologic connectivity influence short-term responses of north temperate lakes to wildfire. Geophysical Research Letters. https://doi.org/10.1029/2023GL103953
McCullough, I. M., Hanly, P. J., King, K. B. S. and T. Wagner. (2022). Freshwater corridors in the conterminous United States: a coarse-filter approach based on lake-stream networks. Ecosphere. https://doi.org/10.1002/ecs2.4326
DePalma-Dow, A., McCullough, I. M. and J. A. Brentrup. (2022). Turning up the heat: long-term water quality responses to wildfires and climate change in a hypereutrophic lake. Ecosphere. https://doi.org/10.1002/ecs2.4271
Lofton, M., Brentrup, J., Beck, W., Zwart, J., Bhattacharya, R., Brighenti, L., Burnet, S., McCullough, I.M., Steele, B., Carey, C., Cottingham, K., Dietze, M., Ewing, H., Weathers, K. and S. LaDeau. (2022). Using near-term forecasts and uncertainty partitioning to improve predictions of oligotrophic cyanobacterial density. Ecological Applications. https://doi.org/10.1002/eap.2590
Orr, R., Bui, A., Klope, M., McCullough, I. M., Lee, M., Motta, C. Mayorga, I., Konicek, K. and H. Young. Context-dependent effects of shifting large herbivore assemblages on plant structure and diversity. (2022). Journal of Ecology. https://doi.org/10.1111/1365-2745.13871
McCullough, I. M., Fergus., C. E. and C. Scott. Macrosystems limnology and beyond: reenvisioning the scale of limnology. (2022). Encyclopedia of Inland Waters, 2nd edition. ISBN 9780124095489. https://doi.org/10.1016/B978-0-12-819166-8.00026-8
Cheruvelil, K. S., Soranno, P. A., McCullough, I. M., Webster, K. E., Rodriguez, L. K. and N. J. Smith. LAGOS-US-LOCUS v1.0: Data module of location, identifiers, and their watersheds in the conterminous U.S. (2021). Limnology and Oceanography Letters. https://doi.org/10.1002/lol2.10203.
Soranno, P. A., Cheruveli, K. S., Liu, B., Wang, Q, Tan, P., Zhou, J., King, K. B. S., McCullough, I. M., Stachelek, J., Bartley, M., Filstrup, C. T., Hanks, E. M., Lapierre, J., Lottig, N. R., Schliep, E. M., Wagner T. and K. E. Webster. Ecological prediction at macroscales using big data: Does sampling design matter? (2020). Ecological Applications. https://doi.org/10.1002/eap.2123
Wagner, T., Lottig, N., Bartley, M., Hanks, E., Schliep, E., Wikle, N., King, K., McCullough, I. M., Stachelek, J., Cheruvelil, K., Filstrup, C., Lapierre, J., Liu, B., Smith, N., Soranno, P., Tan, P., Wang, Q., Webster, K. and J. Zhou. (2020). Increasing accuracy of lake nutrient predictions in thousands of lakes by leveraging water clarity data. Limnology and Oceanography Letters. https://doi.org/10.1002/lol2.10134
Poisson, A. C., McCullough, I. M., Cheruvelil, K. S., Elliot, K. C., Latimore, J. A. and P. A. Soranno. (2020). Quantifying the contribution of citizen science to broad-scale ecological databases. Frontiers in Ecology and the Environment. https://doi.org/10.1002/fee.2128
McCullough, I. M., Skaff, N. K., Soranno, P. A. and K. S. Cheruvelil. (2019). No lake left behind: how well do US protected areas meet lake conservation targets? Limnology and Oceanography Letters, 4(6): 183-192. https://doi.org/10.1002/lol2.10123
Filstrup, C. T., King, K. B. S. and I. M. McCullough. (2019). Evenness effects mask richness effects on ecosystem functioning at macro-scales in lakes. Ecology Letters, 22(12): 2120-2129. https://doi.org/10.1111/ele.13407
McCullough, I. M., King, K. B. S., Stachelek, J., Diaz, J., Soranno, P. A. and K. S. Cheruvelil. (2019). Applying the patch-matrix model to lakes: a connectivity-based conservation framework. Landscape Ecology, 34(11): 2703-2718. https://doi.org/10.1007/s10980-019-00915-7.
McCullough, I. M., Cheruvelil, K. S., Lapierre, J., Lottig, N. R., Moritz, M. A., Stachelek, J. and P. A. Soranno. (2019). Do lakes feel the burn? Ecological consequences of increasing exposure of lakes to fire in the continental United States. Global Change Biology, 25(9): 2841-2854. https://doi.org/10.1111/gcb.14732.
Davis, F. W., Synes, N. W., Fricker, G. A., McCullough, I. M., Serra-Diaz, S. M., Franklin, J. and A. Flint. (2019). LiDAR-derived topography and forest structure predict fine-scale variation in daily surface temperatures in oak savanna and conifer forest landscapes. Agricultural and Forest Meteorology, 269: 192-202. https://doi.org/10.1016/j.agrformet.2019.02.015.
McCullough, I. M., Cheruvelil, K. S., Collins, S. C. and P. A. Soranno. Geographic patterns of the climate sensitivity of lakes. (2019). Ecological Applications, e01836. https://doi.org/10.1002/eap.1836.
McCullough, I. M., Dugan, H. A., Farrell, K. J., Morales-Williams, A. M., Ouyang, Z., Roberts, D. C., Scordo, F., Bartlett, S. L., Burke, S. M., Doubek, J. P., Krivak-Tetley, F. E., Skaff, N. K., Summers, J. C., Weathers, K. C. and P. C. Hanson. (2018). Dynamic modeling of organic carbon fates in lake ecosystems. Ecological Modelling, 386: 71-82. https://doi.org/10.1016/j.ecolmodel.2018.08.009.
Engel, F., Farrell, K. J., McCullough, I. M., Scordo, F., Denfeld, B. A., Dugan, H. A., de Eyto, E., Hanson, P. C., McClure, R. P., Noges, P., Noges, T., Ryder, E., Weathers, K. C. and G. A. Weyhenmeyer. (2018). A lake classification concept for a more accurate global estimate of the dissolved inorganic carbon export from terrestrial ecosystems to inland waters. The Science of Nature, 105:25. https://doi.org/10.1007/s00114-018-1547-z.
McCullough, I. M., Davis, F. W. and A. P. Williams. (2017). A range of possibilities: assessing geographic variation in climate sensitivity of ponderosa pine using tree-rings. Forest Ecology and Management, 402: 223-233. https://doi.org/10.1016/j.foreco.2017.07.025.
Dugan, H. A., Bartlett, S. L., Burke, S. M., Doubek, J. P., Krivak-Tetley, F. E., Skaff, N. K., Summers, J. C., Farrell, K. J., McCullough, I. M., Morales-Williams, A. M., Roberts, D., Scordo, F., Yang, Z., Hanson, P. C. and K. C. Weathers. (2017). Salting our freshwater lakes. Proceedings of the National Academy of Sciences, 114(17): 4453-4458. https://doi.org/10.1073/pnas.1620211114.
Davis, F. W., Sweet, L., Serra-Diaz, J. M., Franklin, J., McCullough, I. M., Flint, A., Flint, L., Dingman, J. R., Regan, H. M., Syphard, A., Hannah, L., Redmond, K. and M. A. Moritz. (2016). Shrinking windows of opportunity for oak seedling establishment in southern California mountains. Ecosphere, 7(11), article e01573. https://doi.org/10.1002/ecs2.1573.
McCullough, I. M., Davis, F. W., Dingman, J. R., Flint, L. E., Flint, A. L., Serra-Diaz, J. M., Syphard, A. D., Moritz, M. A., Hannah, L. and J. Franklin. (2016). High and dry: high elevations disproportionately exposed to regional climate change in Mediterranean-climate landscapes. Landscape Ecology 31(5): 1063-1075. https://doi.org/10.1007/s10980-015-0318-x.
Serra-Diaz, J. M., Franklin, J., Sweet, L, McCullough, I. M., Syphard, A. D., Regan, H., Flint, L., Flint, A., Dingman, J., Moritz, M. A., Redmond, K., Hannah, L. and F. W. Davis. (2016). Averaged 30 year climate change projections mask opportunities for species establishment. Ecography. 844-845. https://doi.org/10.1111/ecog.02074.
Hannah, L., Flint, L. E., Syphard, A. D., Moritz, M. A., Buckley, L. B. and I. M. McCullough. (2015). Place and process in conservation planning for climate change: a reply to Keppel and Wardell-Johnson. Trends in Ecology and Evolution 30(5): 233-234. https://doi.org/10.1016/j.tree.2015.03.008.
Aceves-Bueno, E., Adeleye, A. S., Bradley, D., Brandt, W. T., Callery, P., Feraud, M., Garner, K. L., Gentry, R., Huang, Y., McCullough, I. M., Pearlman, I., Sutherland, S. A., Wilkinson,W., Yang, Y, Zink, T., Anderson, S. E. and C. Tague. (2015). Citizen science as a tool for overcoming insufficient monitoring and inadequate stakeholder buy-in in adaptive management: Criteria and evidence. Ecosystems 18(3): 493-506. https://doi.org/10.1007/s10021-015-9842-4.
Hannah, L., Flint, L., Syphard, A. D., Moritz, M. A., Buckley, L. B. and I. M. McCullough. (2014). Fine-grain modeling of the response of species to climate change: holdouts, stepping-stones and microrefugia. Trends in Ecology and Evolution 29 (7): 390-397. https://doi.org/10.1016/j.tree.2014.04.006.
Dingman, J. R., Sweet, L. C., McCullough, I. M., Davis, F. W., Flint, A., Franklin, J. and L. E. Flint (2013). Cross-scale modeling of surface temperature and seedling establishment to improve projections of tree distribution shifts under climate change. Ecological Processes 2-30. https://doi.org/10.1186/2192-1709-2-30.
McCullough, I. M., Loftin, C. S. and S. A. Sader. (2013). Landsat imagery reveals declining clarity of Maine’s lakes during 1995-2010. Freshwater Science 32(3): 741-752. https://doi.org/10.1899/12-070.1.
McCullough, I. M., Loftin, C. S. and S. A. Sader. (2013). Lakes without Landsat? An alternative approach to remote lake monitoring with MODIS 250 m imagery. Lake and Reservoir Management 29: 89-98. https://doi.org/10.1080/10402381.2013.778926.
McCullough, I. M., Loftin, C. S. and S. A. Sader. (2012). High-frequency remote monitoring of large lakes with MODIS 500 m imagery. Remote Sensing of Environment 124: 234-241. https://doi.org/10.1016/j.rse.2012.05.018.
McCullough, I. M., Loftin, C. S. and S. A. Sader. (2012). Combining lake and watershed characteristics with Landsat TM data for remote estimation of regional lake clarity. Remote Sensing of Environment 123: 109-115. https://doi.org/10.1016/j.rse.2012.03.006.
Technical reports and other publications
Webster, K. E., McCullough, I. M. and P. A. Soranno. (2022). Deeper by the dozen: Diving into a database of 17,675 depths for U.S. lakes and reservoirs. Limnology and Oceanography Bulletin. https://doi.org/10.1002/lob.10482.
McCullough, I. M., Cheruvelil, K. S., Collins, S. C. and P. A. Soranno. Geographic patterns of the climate sensitivity of lakes. Bulletin of the Ecological Society of America: Photo Gallery. Available here.
Moritz, M.A., Topik, C, Allen, C. D., Hessburg, P. F., Morgan, P., Odion, D. C., Veblen, T. T. and McCullough, I. M. (2018). A Statement of Common Ground Regarding the Role of Wildfire in Forested Landscapes of the Western United States. Fire Research Consensus Working Group Final Report. https://www.nceas.ucsb.edu/featured/fire-research-consensus.
McCullough, I. M., Loftin, C. S. and S. A. Sader. A manual for remote sensing of Maine lake clarity. (2013). Maine Agricultural and Forest Experimentation Station, University of Maine. Technical Bulletin 207. ISSN: 1070-1054.